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Folia Primatologica

International Journal of Primatology - Internationale Zeitschrift für Primatologie - Journal international de Primatologie

Guidelines for Authors

Ethical Guidelines
Guidelines for Zoological Nomenclature
Plagiarism Policy
Color Illustrations
References (Note Change in Format)
Digital Object Identifier (DOI)
Supplementary Material
Self-Archiving/Green Open Access, Author's ChoiceTM,
   Funding Organizations (NIH etc.)



Submissions are accepted from any area of primatology (including, for example, behaviour, morphology, physiology, genetics, conservation, evolutionary biology or palaeontology). Our overall criteria for publication are quality and whether papers are likely to be of interest to a general primatological audience. Four types of manuscripts will be considered for publication: Original Research Articles, Reviews (following consultation with the Editors), Technical Reports, and Short Communications for rapid dissemination. Original Research Articles should not normally exceed 10,000 words. For Reviews, we invite experienced researchers to submit brief proposals. Following agreement with the author on a submission and publication schedule, accepted proposals will be published free access and free of charge if the schedule has been adhered to. Reviews should not normally exceed 13,000 words. Technical Reports covering new techniques or reviews of methodology are acceptable if of interest to a broad primatological audience and should not normally exceed 5,000 words. Short Communications of a time-sensitive nature can be fast-tracked for publication, with a target time of 1 month between submission and online publication. News Items should not exceed 3,000 words and their submission must be accompanied by a clear justification for the time sensitivity of their content. All word counts are inclusive of references, tables, figure legends, etc. Longer manuscripts may exceptionally be accepted for publication, but the Editors reserve the right to insist that any manuscript be shortened if they feel that the length is not justified. All types of manuscripts are subjected to full peer review. Papers of preliminary or anecdotal content are unlikely to be reviewed unless they are of unusual and/or immediate interest, such as techniques of high potential for primatological research or discoveries of new species. Proposals for themed Special Issues are welcome and should be submitted to the Editors.

Manuscripts should be written in International English. An abstract in English must be included.

Names, postal and email addresses of four potential referees with expertise relevant to the research presented in the manuscript should be included with the submission. Referees suggested should not be from the same institution as the author(s) nor be research collaborators of the author(s).

Manuscripts should be submitted online.

Should you experience any problems with your submission, please contact:

Lara Grieder
Editorial Office ‘Folia Primatologica’
S. Karger AG
P.O. Box
CH–4009 Basel (Switzerland)
Tel. +41 61 306 1356
Fax +41 61 306 1434

Do not send manuscripts or revised manuscripts to any other address. Do not address queries concerning progress to any other address.

In an effort to reduce turnaround times and in fairness to other authors, authors are requested to return revised manuscripts within two months of receipt of the reviewers’ comments and the editor’s decision letter (one week for fast-tracked Short Communications). If this is not possible, it is essential for authors to notify the Editorial Office in writing of any delay, with reasons for the delay, within these time limits. Except where prior agreement was reached to extend the deadline, manuscripts that are returned after the deadline will be treated as new submissions.

Ethical Guidelines

Folia Primatologica requires that all papers reporting field studies in which animals have been trapped, darted or otherwise handled, and papers reporting studies of captive animals, other than purely observational studies, should include in the text a clear statement of the justification for any animal contact and/or invasive procedures. It is not sufficient to state that protocols involving contact, or invasive procedures, accord to regulations of an internal or national ethical committee.

Folia Primatologica recognises that samples need to be taken to establish type specimens for new taxa, but this can and should be done in such a way as to minimize death or other loss to wild populations: conservation criteria should take priority over the clarification of taxonomic status.

Folia Primatologica will not normally accept papers which rely on taking and killing of multiple wild primates as ‘voucher’ specimens or for any other reason, and any submission involving such research must be agreed with the Editors in advance and be thoroughly justified to them in animal/human welfare or conservation terms.

When animals are euthanized during or at the end of experiments, justification in terms of a clear benefit to the health or well-being of humans and/or non-human primates is essential, even in conference abstracts.

When cadavers of wild primates, or live primates are bought from markets or local dealers for use in research, 'Folia Primatologica' expects a statement to be made indicating how it was ensured that such activity would not result in increased hunting/killing or collecting by the local community.

The Editors' decision on ethical issues is final.

Guidelines for Zoological Nomenclature

'Folia Primatologica' requires that authors adhere exactly to the provisions of the International Code For Zoological Nomenclature. This is particularly important when making claims for the existence of a new taxon.


All manuscripts are subject to editorial review.

Manuscripts are received with the explicit understanding that they are not under simultaneous consideration by any other publication. Submission of an article for publication implies the transfer of the copyright from the author to the publisher upon acceptance.

Accepted papers become the permanent property of 'Folia Primatologica' and may not be reproduced by any means, in whole or in part, without the written consent of the publisher.

For legal reasons, we must receive your 'Submission Statement' with your original (hand-written) signature. Please download, print, sign and either fax or scan it to make it legally binding.

It is the author's responsibility to obtain permission to reproduce illustrations, tables, etc. from other publications.

Plagiarism Policy

Whether intentional or not, plagiarism is a serious violation. We define plagiarism as a case in which a paper reproduces another work with at least 25% similarity and without citation.

If evidence of plagiarism is found before/after acceptance or after publication of the paper, the author will be offered a chance for rebuttal. If the arguments are not found to be satisfactory, the manuscript will be retracted and the author sanctioned from publishing papers for a period to be determined by the responsible Editor(s).


Lines should be numbered on the margin.

Title page: The first page of each paper should indicate the title, the authors' names, the institute where the work was conducted, and a short title for use as running head. A count of the number of words in the manuscript, including main text, references and legends for figures and tables must be given.

Full address: The exact postal address of the corresponding author complete with postal code must be given at the bottom of the title page. Please also supply phone and fax numbers, as well as e-mail address.

Key words: For indexing purposes, a list of 6–10 key words in English is essential.

Abstract: Each paper needs an abstract of up to 200 words in English.

Footnotes: Avoid footnotes as far as possible. When essential, they are numbered consecutively and typed at the foot of the appropriate page.

Tables and illustrations: Tables and illustrations (both numbered in Arabic numerals) should be prepared on separate sheets. Tables require a heading and figures a legend, also prepared on a separate sheet. For the reproduction of illustrations, only good drawings and original photographs can be accepted; negatives or photocopies cannot be used. Due to technical reasons, figures with a screen background should not be submitted. When possible, group several illustrations on one block for reproduction (max. size 144 x 196 mm) or provide crop marks. B/w half-tone and colour illustrations must have a final resolution of 300 dpi after scaling, line drawings one of 800–1,200 dpi.

Colour Illustrations

Online edition: Colour illustrations are reproduced free of charge. In the print version, the illustrations are reproduced in black and white. Please avoid referring to the colours in the text and figure legends.

Print edition: Up to 6 colour illustrations per page can be integrated within the text at CHF 960.00 per page.


In the text, give author’s name and date of publication in brackets, with et al. for three or more authors. Multiple references should be listed chronologically. The list of references should be arranged alphabetically (except that two-authored papers are listed before multi-authored papers with the same first author) and should include only those publications cited in the text. Work accepted for publication may be listed as [in press],[unpublished observations] or [personal commu­nications] cited in the text should not be listed in the references.

Please punctuate and format references exactly as in the following examples.

Papers published in periodicals:
DelPero M, Pozzi L, Masters JC (2006). A composite molecular phylogeny of living lemuroid primates. Folia Primatologica 77: 434–445.

Papers published only with DOI numbers:
Baden AL, Holmes SM, Johnson SE, Engberg SE, Louis EE Jr, Bradley BJ (2014). Species-level view of population structure and gene flow for a critically endangered primate (Varecia variegata). Ecology and Evolution DOI: 10.1002/ece3.1119.

Swindler DR (2002). Primate Dentition: An Introduction to the Teeth of Non-Human Primates. Cambridge, Cambridge University Press.

Edited books:
Di Fiore A, Campbell CJ (2007). The atelines: variation in ecology, behaviour and social organization. In Primates in Perspective (Campbell CJ, Fuentes A, Mackinnon KC, Panger M, Bearder SK, eds.), pp 155–185. New York, Oxford University Press.

Digital Object Identifier (DOI)

S. Karger Publishers supports DOIs as unique identifiers for articles. A DOI number will be printed on the title page of each article. DOIs can be useful in the future for identifying and citing articles published online without volume or issue information. More information can be found at www.doi.org

Supplementary Material

Multimedia files and other supplementary files, directly relevant but not essential to the conclusions of a paper, enhance the online version of a publication and increase its visibility on the web. These files will undergo editorial review. The Editors reserve the right to limit the scope and length of the supplementary material. Multimedia and supplementary material should meet production quality standards for publication without the need for any modification or editing. Files should not exceed 10 MB in size. Figures and tables need to have titles and legends, and all files should be supplied separately and labeled clearly. All supplementary material should be referred to in the main text. A DOI number will be assigned to supplementary material and it will be hosted online at https://karger.figshare.com under a CC BY license. Authors will be charged a processing fee of CHF 250.00 for supplementary material.

Self-Archiving/Green Open Access

Karger permits authors to archive their pre-prints (i.e. pre-peer review) or post-prints (i.e. accepted manuscript after peer review but before production) on their personal or their institution’s internal website. In addition, authors may post their accepted manuscripts in public Open Access repositories and scientific networks (e.g. ResearchGate or Mendeley) no earlier than 12 months following publication of the final version of their article. For all self-archiving, the posted manuscripts must:

  • Be used for noncommercial purposes only
  • Be linked to the final version on www.karger.com
  • Include the following statement:

‘This is the peer-reviewed but unedited manuscript version of the following article: [insert full citation, e.g. Cytogenet Genome Res 2014;142:227–238 (DOI: 10.1159/000361001)]. The final, published version is available at http://www.karger.com/?doi=[insert DOI number].’

It is the author’s responsibility to fulfill these requirements.

For papers published online first with a DOI number only, full citation details must be added as soon as the paper is published in its final version. This is important to ensure that citations can be credited to the article.

Manuscripts to be archived in PubMed Central due to funding requirements will be submitted by Karger on the author’s behalf [see Funding Organizations (NIH etc.)].

For self-archiving Author's ChoiceTM (Gold Open Access) articles, see Author's ChoiceTM.

Author's ChoiceTM

Karger’s Author's ChoiceTM service broadens the reach of your article and gives all users worldwide free and full access for reading, downloading and printing at www.karger.com. The option is available for a one-time fee of CHF 3,000.00, which is a permissible cost in grant allocation. More information can be found at www.karger.com/authors_choice.
The final, published version of the article may be posted at any time and in any repository or on other websites, in accordance with the relevant Creative Commons license. Reposted Open Access articles must:

  • Follow the terms of the relevant Creative Commons license
  • Be linked to the final version on www.karger.com
  • Include the following statement:

‘The final, published version of this article is available at http://www.karger.com/?doi=[insert DOI number].’

It is the author’s responsibility to fulfill these requirements.

For papers published online first with a DOI number only, full citation details must be added as soon as the paper is published in its final version. This is important to ensure that citations can be credited to the article.

Funding Organizations (NIH etc.)

The U.S. National Institutes of Health (NIH) Public Access Policy mandates that accepted, peer-reviewed manuscripts are archived in its digital database, PubMed Central (PMC), within 12 months of the official publication date. As a service to authors, Karger submits NIH-funded articles to PMC on behalf of the authors immediately upon publication. The NIH assigns a PMCID within approximately 1 month and the manuscript will appear in PMC after a 12-month embargo. For authors making their paper Open Access through Author's ChoiceTM, the embargo will be overridden, thereby accelerating the accessibility of the article. Karger also complies with other funders’ requirements (including Wellcome Trust and RCUK) for submission to PMC.
Authors should include information on their grant in the Acknowledgements section of their papers.


Unless indicated otherwise, proofs are sent to the corresponding author and should be returned with the least possible delay. Alterations made in proofs, other than the correction of printer's errors, are charged to the author.


Order forms and a price list are sent with the proofs. Orders submitted after the issue is printed are subject to considerably higher prices.